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GHK-Cu research themes may include: collagen marker research fibroblast activity extracellular matrix research matrix metalloproteinase activity glycosaminoglycan marker studies skin model research copper peptide biology tissue remodelling markers Another PubMed-indexed study reported that GHK-Cu stimulated matrix metalloproteinase-2 expression in fibroblast cultures, supporting its role in extracellular matrix remodelling research